Leonardo da Vinci described the heart as vortex-oriented around 1512–1513, suggesting that blood flows in spiral patterns through the ventricles and forms vortices in the aortic sinuses, which are essential for closing the aortic valve, rather than flowing in a simple, straight line. Modern 4D-MRI and fluid modeling techniques have confirmed these predictions 500 years later, proving that the heart acts as a biological vortex generator, improving efficiency and reducing energy loss.
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